Perona

A girl with the power of the Hollow-Hollow Fruit who was once a member of the Thriller Bark Pirates. Known as the "Ghost Princess," Perona made her major debut in *One Piece* as one of the key commanders of the Mysterious Four aboard the ship-island Thriller Bark, serving the former Shichibukai Gecko Moria. Brought into the crew at a young age, she grew up surrounded by bizarre creatures and zombies, forging a strong bond of loyalty with Moria and taking command of the location's servants and wild zombies. Her path changed drastically during the Straw Hats' invasion of Thriller Bark, where she was defeated in an unusual manner by Usopp. While attempting to flee the ship amidst the chaos, Perona crossed paths with Bartholomew Kuma, who used his powers to send her to the remote Kuraigana Island. There, she lived alongside Dracule Mihawk and Roronoa Zoro during the two-year timeskip, establishing a comical dynamic of bickering and care before helping Zoro reunite with his friends and setting out to discover Moria's whereabouts. Perona's personality is a striking blend of childishness, vanity, and bossiness, all wrapped in a style inspired by Gothic Lolita fashion. She is deeply attached to things she considers cute—though her definition of "cute" includes terrifying or macabre creatures that would frighten ordinary people. Despite her often spoiled, bossy, and temperamental behavior, she possesses an empathetic and caring side, evident when she tended to Zoro's severe injuries without hesitation. In combat, her abilities rely on the *Horo Horo no Mi* (Hollow-Hollow Fruit), a Paramecia-type Devil Fruit that allows her to generate and manipulate ghosts. Her most famous and devastating ability is "Negative Hollow," in which she releases spirits that pass through opponents' bodies, instantly draining their self-esteem and will to fight, leaving even the proudest warriors prostrate and overcome by absolute pessimism. Beyond draining enemies' morale, Perona can create ghosts of various sizes that explode on her command ("Toku Hollow"), inflicting real physical impact. She can also use astral projection to separate her soul from her physical body, becoming intangible and capable of flight to spy on distant areas or alter her size, though her actual body must remain protected and motionless during the process. The major limitation of her negative ghosts is that they have no effect on individuals who are already naturally pessimistic and lack self-confidence. Even without relying on brute physical strength, the combination of mind-control powers, tactical versatility, and a striking appearance has made Perona an unforgettable figure. Her evolution from an eccentric antagonist into a loyal and charismatic supporting character demonstrates how she has established herself as one of the most unique and beloved figures in the series' universe.